Thomas C. Harrington. Research on the evolution and ecology of forest pathogens and other fungi, including Ceratocystis, Ophiostoma, Raffaelea, Tubakia, Heterobasidion and Armillaria, bluestain, bark beetle fungi, diseases of poplar and soybean, and the role of disease in forest ecosystems.

Iowa State University, Department of Plant Pathology
221 Bessey Hall, Ames, Iowa 50011
Office: 515-294-0582, FAX: 515-294-9420


Tom Harrington received his B.S. Degree in Plant Pathology from Colorado State University in 1977, his M.S. in Plant Pathology from Washington State University in 1980, and his Ph.D. in Plant Pathology from the University of California at Berkeley in 1983. He was on the faculty of the University of New Hampshire from 1983 through 1990. mango

 He has been a member of the

ISU Plant Pathology Department since 1991 and has had a joint appointment in the

ISU Forestry Department since 1996. The Forestry Department merged with Animal Ecology to form Natural Resource Ecology and Management Department .

He is also a member of the Microbiology major .

 Click on the links above to see the graduate programs in which my students can major.
For photographs of people, places, plants:
People (Updated April 2014)
Cacao History

Descriptions, Publications, and Images Concerning Specific

Research Projects Can be Found by Clicking the Topic Below

 Ceratocystis and other Ceratocystidaceae, including Ambrosiella

Bur oak blight (BOB), caused by Tubakia iowensis

 Ophiostoma, Raffaelea, Leptographium, Sphaeropsis, Amylostereum:

Bark and ambrosia beetle fungi, bluestain fungi, horntail fungi, and other insect-associated fungi

Also, a special page for laurel wilt

 Species Concepts, Canopy Gaps, Forest Ecology, Wind, Cryphonectria, Techniques and Crops

Poplar Diseases (microsatellites for Melampsora spp.)

 Heterobasidion, Armillaria, and Other Root and Butt Rot Fungi

 Soorty blotch/Flyspeck, Sclerotium, Cercospora, Cadophora, Boremia, Phialophora, Phytoplasmas, and Soybean Diseases

Video links, Illustrations and Downloadable Book Chapters:

Bur oak blight - caused by a new Tubakia species:

A recently published review of the disease in Diseases of the Great Plains. (pdf of book chapter)

Distribution and intensification of bur oak blight in Iowa and the Midwest. (pdf of book chapter).

The formal description of bur oak blight and the new species (Tubakia iowensis) is available in pdf form.

This is a link to a 2010 video (actually a powerpoint presentation) on bur oak blight - "What about BOB?"

A recent article (Feb. 2011) on bur oak blight was published in the ISU Extension Horticulture & Home Pest Newsletter.

The US Forest Service has a 2-page pest alert on bur oak blight.

 Also available in PDF format: T. C. Harrington. 2005.

Ecology and Evolution of Mycophagous Bark Beetles and Their Fungal Partners

In: Insect-Fungal Associations. Ecology and Evolution,

F. E. Vega and M. Blackwell, eds. Oxford University Press

Illustrations of various
Diseases caused by Ceratocystis
can be found through this link. Diseases of cacao, inhame, sweet potato, mango, coffee, aspen, sycamore, hickory, almond, maple, Nothofagus, palms, and oak are illustrated thus far.
Our work on Ceratocystis species was supported bythe National Science Foundation (DEB-9870675 and DEB-0128104).

 A review of the liturature on

Ceratocystis fimbriata

and the diseases caused by this complex of species, initially prepared for CABI's 2001 Crop Protection Compendium, revised for the 2004 compendium, and updated frequently on this site:




Ceratocystis wilt of cacao
(Theobroma cacao, the tree that gives us chocolate) has been added. This gives more detailed information on Ceratocystis fimbriata in cacao, especially in Bahia, Brazil.

 Images of:

Diseases and pine ecology

can be found here. They go with the book chapter to the right:

 Available in PDF format: Harrington, T. C., and M. J. Wingfield. 1998.

Diseases and the ecology of indigenous and exotic pines.

pp. 381-404 In: D. Richardson, ed. Ecology and Biogeography of Pinus. Cambridge University Press.Simply click on icon to download: pdf image

To find out more about the book, check with Cambridge University Press.

 Also available in PDF format: Harrington, T. C., and D. M. Rizzo. 1999.

Defining species in the fungi.

pp. 43-71 In: J. J. Worrall, ed., Structure and Dynamics of Fungal Populations, Kluwer Press. Dordrecht, Netherlands. Simply click on icon to download: pdf image

 Also available in PDF format: Paine, T. D., K. F. Raffa, and T. C. Harrington. 1997.

Interactions among scolytid bark beetles, their associated fungi, and live host conifers.

Annual Rev. Entomol. 42:179-206. Simply click on icon to download: pdf image

There is a great web site on the Trees of the ISU Campus!

ISU Plant Pathology Department
Mycological Society of America Homepage
This page updated April 23, 2014

eXTReMe Tracker